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Role of how to articles College plant glyoxylate reductases during stress: a hypothesis. Molecular modelling suggests that a group of proteins in plants known as the ?-hydroxyacid dehydrogenases, or the hydroxyisobutyrate dehydrogenase superfamily, includes enzymes that reduce succinic semialdehyde and glyoxylate to ?-hydroxybutyrate and introduction for persuasive School, glycolate respectively. How To Articles Seneca. Recent biochemical and expression studies reveal that NADPH-dependent cytosolic (termed GLYR1) and plastidial (termed GLYR2) isoforms of succinic semialdehyde/glyoxylate reductase exist in Arabidopsis . Succinic semialdehyde and glyoxylate are typically generated in leaves via two distinct metabolic pathways, ?-aminobutyrate and glycolate respectively. Writing Scientific School. In the present review, it is proposed that the GLYRs function in the detoxification of both aldehydes during stress and contribute to redox balance. Outstanding questions are highlighted in a scheme for the subcellular organization of the how to write articles detoxification mechanism in Arabidopsis . aldehyde detoxification glyoxylate reductase ?-hydroxyacid dehydrogenase redox balance succinic semialdehyde reductase. The occurrence of Warminster distinct NADPH-dependent GLYR (glyoxylate reductase) isoforms and activity in the cytosol and write feature articles Seneca, chloroplasts of spinach and pea has been documented . Recent research has indicated that Arabidopsis cytosolic GLYR1 and plastidial GLYR2 reduce both glyoxylate and info School Eerde, SSA (succinic semialdehyde) to glyoxylate and GHB (?-hydroxybutyrate) respectively [2–4]. In vitro measurements of recombinant GLYR1 and GLYR2 activities revealed that glyoxylate is preferred over SSA as a substrate, the kinetic mechanism is ordered Bi Bi, involving the complexation of how to write articles Seneca College NADPH before glyoxylate or SSA, and NADP is a competitive inhibitor with respect to NADPH. Thus plant GLYRs act as an AKR (aldo-keto reductase), but there is no significant homology with either mammalian and bacterial NADPH-SSA reductase, or other members of the AKR superfamily. There is, however, some sequence identity with known ?-hydroxyacid dehydrogenases, such as tartronate semialdehyde reductase and 6-phosphogluconate dehydrogenase, which are members of the HIBADH (hydroxyisobutyrate dehydrogenase) protein family. Furthermore, molecular modelling of GLYR1 via comparison with a well-characterized member of the HIBADH family indicates that a Rossmann fold-like nucleotide-binding site with a three-layer (???) sandwich geometry at the N-terminus, orthogonal bundles at the C-terminus, and amino acid residues at the predicted active site are highly conserved (Figure 1).
Taken together, these observations suggest that plant GLYR enzymes are members of the HIBADH protein family [2,3], but crystallization of the proteins and active-site mutagenesis are necessary to substantiate this hypothesis. ( A ) Crystal structure of TSAR (tartronic semialdehyde reductase) and its substrate analogue tartaric acid (TA) (PDB entry: 1VPD). Introduction For Persuasive Essay Warminster School. ( B ) Predicted structure of GLYR1 generated by the fold-recognition server PHYRE (http://www.sbg.bio.ic.ac.uk/phyre/) using TSAR as a template. ( C and D ) Overlapping positions and identities of the predicted active-site residues for TSAR and GLYR1 respectively. Red, blue and how to feature, green indicate negative, positive and hydrophobic regions of the residues. Aldehydes such as glyoxylate and SSA can accumulate in plants under stress, and writing tricks, react with DNA, oxidize membrane lipids, modify proteins or influence the transcription of stress-related genes, thereby causing cellular and developmental problems [5,6]. Consequently, enzymes and metabolic pathways that reduce the aldehyde chemical grouping (i.e. H-C=O) in these compounds to its corresponding alcohol are probably essential for maintaining plant health. Glyoxylate and how to write articles Seneca College, SSA are best known as intermediates in the metabolism of School glycolate and GABA (?-aminobutyrate) respectively. In the present review, we attempt to place the function of how to write feature articles College plant GLYRs within a metabolic context. The first three sections discuss the physiological role of the introduction for persuasive essay Warminster School enzymes during stress, with particular emphasis on feature articles College the turnover of pyridine nucleotides and the metabolism of GABA and written King’s Ely, glycolate. The final section focuses on Seneca a hypothetical scheme for the detoxification of SSA and introduction essay School, glyoxylate, and on opportunities for future research. NAD(P) is ubiquitous in the cell, mediates numerous redox reactions, and how to articles, influences virtually every metabolic pathway [7–9].
Furthermore, it is writing articles School, involved in the production and consumption of ROS (reactive oxygen species) in plants, and is therefore a key player in signalling events. ROS production is influenced by NADPH/NADP ratios in chloroplasts and mitochondria and NAD(P)H oxidases are key players in how to feature Seneca ROS generation at the plasma membrane. ROS consumption partly depends on glutathione and ascorbate pools, which are fundamentally maintained by NAD(P)H. Writing Scientific Articles Pomfret. NAD(P) plays other defensive and signalling roles, such as production of NO and metabolism of reactive aldehydes. NAD(P) pools are plastic in feature articles Seneca plants and undergo significant changes in response to the environment. For example, increasing light causes progressive over-reduction at the reducing side of Photosystem I in the chloroplast, causing NADP levels to decline and thereby favouring the diversion of electrons to oxygen to Nuova Accademia Milano produce ROS . How To Articles College. Yet the chloroplast stromal redox state seems to info International School be constant in spite of write feature articles irradiance. The cytosolic NAD(P) redox states are independent of scientific articles Pomfret light, dark or CO 2 availability, yet cytosolic NAD and mitochondrial NAD and NADP are more reduced under photorespiratory conditions . Several other studies indicate that changes in how to write articles Seneca College cytosolic and mitochondrial NAD reduction are important in regulating nitrate reduction and nitrogen assimilation [12–14].
Generation of NADPH in writing School the chloroplast depends on the reduction of NADP via ferredoxin NADP reductase in the light, and by the oxidative pentose phosphate pathway dehydrogenases, glucose-6-phosphate dehydrogenase and 6-phosphogluconate dehydrogenase, in the dark [10,14]. The presence of glucose-6-phosphate and 6-phosphogluconate dehydrogenases in peroxisomes implies that these organelles have the write feature articles Seneca capacity to custom written essays King’s Ely reduce NADP to NADPH for use in antioxidant protection . In the cytosol and mitochondria, NADP-dependent ICDHs (isocitrate dehydrogenases) play a major role in NADPH pool sizes [11,16]. NADP-ICDH activity in mitochondria is higher in photosynthetic than in heterotrophic tissues, implying an important role during photosynthesis. It has been suggested that mitochondrial NADP-ICDH provides NADPH for redox regulation of the alternative oxidase via thioredoxin . The NADH/NAD ratio in mitochondria is high during photorespiratory conditions due to glycine oxidation via GDC (glycine decarboxylase) activity, which provides more NADH than the TCA cycle (tricarboxylic acid cycle; also known as the articles Seneca College citric acid cycle) [11,18]. This higher ratio could stimulate the interconversion of NADH and NADP via transhydrogenase activity [11,19], thereby increasing mitochondrial NADPH levels, inhibiting NAD-ICDH in the TCA cycle, and essay American School, causing mitochondrial NADP-ICDH to how to articles operate in the reverse direction. Consequently, citrate could be exported from the mitochondrion, whereas 2-oxoglutarate could be imported, entering the reverse NADP-ICDH reaction as 2-oxoglutarate dehydrogenase activity is probably low during photorespiration.
Thus NADP-ICDH reverse activity might recycle NADPH and 2-oxoglutarate in the mitochondrion during photorespiration. Mitochondrial NADP and NAD pools are more oxidized in the dark , and the NADH/NAD ratio is not significantly lower than under non-photorespiratory conditions, which represents oxidation of NADH mainly through the TCA cycle and not through GDC. The ATP-dependent phosphorylation of NAD(H) is catalysed by NADK (NAD kinase). Genes encoding NADKs have been cloned from human, yeast and bacteria [20–22]. Eukaryotes possess multiple NAD(H)K genes, whereas bacteria apparently have only written, one. The enzymes are divided into NADKs or NAD(H)Ks depending on their substrate preference for how to feature the oxidized and/or reduced forms of NAD [23,24]. Several NADK isoforms are present in introduction essay Warminster School plants. In maize, for example, a CaM (calmodulin)-dependent enzyme is found in the cytosol, in College the outer membrane of mitochondria and in the chloroplast envelope, whereas a CaM-independent form is located in the chloroplast stroma [25,26]. In Avena sativa L. Writing Pomfret School. seeds, soluble and mitochondrial NADK activities are present in CaM-dependent and -independent forms [27,28]. CaM-dependent activity has also been detected in pea seedlings, Solanum lycopersicum L. var. pimpinellifolium (Juslen) Mill and green bean [29,30].
In Arabidopsis , there are three isoforms: two are cytosolic [ At NADK1 and At NAD(H)K3] and one is plastidial ( At NADK2) [31–34]. Partially purified NADK activity from how to feature articles College, Arabidopsis displays Ca 2+ /CaM-dependent activity, whereas recombinant At NAK1 and At NADK2 do not, although At NADK2 has some affinity for CaM . One study has reported that recombinant At NADK1 can utilize both NAD and written King’s, NADH in an ATP-dependent manner, although the substrate specificity constant was not determined , whereas another reported that At NADK1 and At NADK2 utilize NAD, but not NADH . At NAD(H)K3 is highly homologous with POS5, a mitochondrial isoform from yeast recently described as an NADK, which prefers NAD(H) as a substrate, but it has negligible amino acid identity with other plant NADKs [23,31]. At NAD(H)K3 can phosphorylate NADH or NAD + in an ATP-dependent manner, but has a 100-fold greater preference for NADH, and its activity is not stimulated by CaM . At NADK1 is write articles, induced by oxidative stress, such as ionizing radiation and paraquat treatment, and by compatible and incompatible plant–pathogen interactions , and coursework info School, NAD(H)K3 is induced by a variety of write articles Seneca College abiotic stresses and exogenous abscisic acid. Nuova Accademia Di Belle Arti. NAD(H)K3 apparently plays an how to feature Seneca College important role in cytosolic NADPH levels under abiotic stress .
Manipulation of At NADK2 levels affects chloroplastic NADPH levels, and null mutants are stunted, with a pale yellow colour, and are hypersensitive to abiotic stress . SSA IS AN INTERMEDIATE IN GABA METABOLISM. Glutamate can be metabolized via a short metabolic pathway that bypasses the research paper about Warminster School NAD-dependent 2-oxoglutarate dehydrogenase and the ADP-dependent succinyl-CoA ligase steps of the how to write feature articles Seneca College TCA cycle and allows the glutamate carbon backbone to enter the info International Eerde cycle as succinate [35,36] (Figure 2). This pathway is found in virtually all organisms from bacteria to humans, underscoring its fundamental nature in metabolism. GABA is a four-carbon non-protein amino acid with the amino group on the ?-carbon, rather than the how to write feature articles Seneca College ?-carbon. The decarboxylation of research about glutamate is catalysed via a cytosolic Ca 2+ /CaM-dependent GAD (glutamate decarboxylase) with the consumption of a proton . The catabolism of GABA apparently involves mitochondrial pyruvate- and Seneca, glyoxylate-dependent, but not 2-oxoglutarate-dependent, GABA-T (GABA transaminase) activities [37–39], resulting in the production of coursework info International Eerde alanine and glycine respectively. Articles Seneca. GABA-T activity also produces SSA which is quickly oxidized via mitochondrial NAD-dependent SSADH (SSA dehydrogenase) [40–42]. (GAD, GABA-T and SSADH make up the pathway that is custom written essays, commonly known as the GABA shunt.) The GABA shunt metabolites and enzymes are indicated as blue and how to Seneca, orange balloons respectively, and the TCA cycle is writing articles, shown as black and pink balloons. Glutamate, the precursor to GABA, is supplied predominantly via the glutamine synthetase/glutamate synthase cycle or the TCA cycle. The polyamine putrescine has been proposed as an how to write feature articles alternative GABA precursor, although in planta contribution in mature leaves is likely to be minor.
There are two branching routes for for persuasive essay Warminster School GABA catabolism via SSA: one to how to write feature succinate and the TCA cycle and the other to GHB via SSA reductase (a yellow balloon, shown as GLYR in Figure 3), although the products of GHB catabolism in plants are currently unknown. The GABA shunt provides metabolic flexibility to the cell with the provision of glutamate rather than pyruvate carbon for use in the TCA cycle. The oxidation of glutamate is theoretically less energetically productive than pyruvate, however, because 1 mol each of essay Leysin School NADH and ATP are produced at the 2-oxoglutarate dehydrogenase and how to write feature Seneca College, succinyl-CoA ligase steps respectively, whereas only 1 mol of NADH is produced at the SSADH step in the GABA shunt. Nevertheless, stresses that restrict the oxidation of NADH, diminish TCA cycle intermediates and/or inhibit TCA cycle enzymes should permit greater flux through the GABA shunt. Articles School. For example, oxygen deficiency inhibits NADH oxidation via the mitochondrial electron transport chain, decreases the glutamate level, and increases the GABA and alanine levels [43–47]. Write Articles College. Furthermore, manipulation of TCA cycle enzymes, such as NAD-dependent ICDH or aconitase, alters GABA shunt activity [48,49], and GABA shunt activity can complement reduced expression of succinyl-CoA ligase .
GABA accumulation in plants is writing Leysin American School, a general response to various biotic and write articles College, abiotic stresses and articles School, can involve both biochemical and transcriptional processes [35,36,50]. How To Feature Seneca. In some cases, the response of GABA is rapid, often within seconds [51,52], suggesting that activation of cytosolic GAD activity by Ca 2+ /CaM is probably involved. It is known that Ca 2+ influx into the cell is increased by touch , cold shock , heat shock, salinity, drought and osmotic stress , and GABA does accumulate. However, at acidic pH GAD is stimulated independently of Ca 2+ /CaM, and in vivo cytosolic acidification does stimulate GAD activity and GABA production [55,56]. Stresses that disrupt cellular membranes such as wounding, freezing or heat stress, can cause vacuolar contents to introduction for persuasive Warminster be released into the cytosol or inhibit ATPase enzymes which pump protons from the cytosol into the vacuole or apoplast, causing a concomitant decrease in cytosolic pH and stimulation of GAD activity. For example, oxygen deficiency is write feature Seneca, known to decrease cytosolic pH by 0.3–0.6 pH units and cause GABA to accumulate in soybean, tobacco and Arabidopsis [43,44,47,57]. Transcriptional change in GAD and GABA-T expression can occur in response to low oxygen, water deficit, salinity or Agrobacterium infection [47,58–61]. Evidence is available for about Warminster the reduction of SSA and accumulation of GHB in Arabidopsis plants subjected to oxygen deficiency, high light, heat, drought and salinity, and for write feature articles College the recovery of GHB levels in tobacco after the essay writing tricks American removal of oxygen deficiency [42,44,46,47,50,62] (Figure 3). In many cases, these results have been associated with altered redox levels. Furthermore, ssadh -knockout mutants of Arabidopsis exhibit chlorotic lesions and accumulate GHB and how to write articles Seneca, H 2 O 2 . GLYR isoforms are also implicated in GHB production because their transcript levels can increase under various stresses, such as cold and heat .
GLYR activity is found at all developmental stages and in all tissues, and activity is generally higher in vegetative and reproductive organs than in roots . SSADH-deficient yeast complemented with Arabidopsis GLYR1 grows on 20 mM GABA as the sole nitrogen source and contains elevated levels of GHB . These studies link the GABA shunt to redox homoeostasis in research paper about Warminster School the cell and suggest that SSA-dependent GLYR activity potentially occurs in how to write feature Seneca planta , despite the fact that glyoxylate is the preferred substrate in vitro . The subcellular scheme relates recent and essays, hypothetical information about GLYRs and NADKs to pyridine nucleotides and specific intermediates of the GABA and glycolate pathways. There is articles Seneca College, no consideration of stoichiometry in this scheme. Enzymes of the GABA and articles, glycolate pathways, as well as photorespiratory nitrogen metabolism, are shown as orange, pink and brown balloons respectively.
Enzymes associated with aldehyde detoxification and Seneca, NADPH formation are shown as blue and turquoise balloons respectively. Introduction Essay Warminster School. Components whose levels are probably elevated in response to stress are shown as yellow boxes, and potential diffusion pathways are shown as dotted lines. For further discussion and references, please see the text. FR, ferredoxin reductase; GGAT, glutamate:glyoxylate aminotransferase; GK, glycerate kinase; GO, glycolate oxidase; GOGAT, glutamate synthase; GS, glutamine synthetase; HPR, hydroxypyruvate reductase; PGP, phosphoglycolate phosphatase; SGAT, serine:glyoxylate aminotransferase; SHMT, serine hydroxymethyltransferase. GLYOXYLATE IS AN INTERMEDIATE IN GLYCOLATE METABOLISM. Rubisco (ribulose-1,5-bisphosphate carboxylase/oxygenase; EC 184.108.40.206) can catalyse the oxygenation of ribulose bisphosphate, resulting in the production of glycolate 2-phosphate, which is metabolized via the glycolate pathway (Figure 3). (This pathway is often referred to collectively as photorespiration.) The rate of oxygenation compared with carboxylation is dependent on the ambient O 2 and CO 2 concentrations, their K m values and the velocity of the overall reaction .
Sharkey et al.  estimated that, under ambient conditions, the rate of photorespiratory CO 2 release is approx. 25% of the how to write rate of net CO 2 assimilation. With rising temperatures, the specificity of Rubisco for CO 2 and the relative solubility of O 2 to info International CO 2 increase, resulting in enhanced photorespiratory rates. This effect is further augmented by a lowered intercellular CO 2 concentration, which occurs upon stomatal closure during drying conditions such as high light and drought , thereby imposing strong carbon loss and possibly leading to the depletion of carbohydrates and accelerated senescence. However, the suppression of photorespiration over the long term results in decreased rates of photosynthetic CO 2 assimilation, poor plant growth and chloroplast aberrations [66,67]. The photorespiratory pathway is compartmentalized across chloroplasts, peroxisomes and Seneca College, mitochondria (Figure 3). Glycolate 2-phosphate is first hydrolysed in King’s Ely the chloroplast to glycolate, which is oxidized in Seneca College the peroxisome to glyoxylate.
The glyoxylate in turn undergoes transamination to glycine [68,69], which is transported to the mitochondrion for conversion into serine via NAD-dependent GDC and serine hydroxymethyltransferase, with the release of CO 2 and NH 3 . Serine returns to the peroxisome where it is transaminated to hydroxypyruvate, which is reduced to glycerate. Glycerate is then phosphorylated in custom King’s Ely the chloroplast and the resulting glycerate 3-phosphate enters the Calvin cycle. How To Write Articles. Because of the transamination of glyoxylate to glycine and the large release of writing scientific Pomfret School NH 3 in the GDC reaction, photorespiratory metabolism is intimately linked to feature articles Seneca College leaf nitrogen metabolism . The energy requirements for fixing CO 2 are higher in review NABA, Nuova Accademia di Belle the presence of photorespiratory metabolism (5.375 mol of ATP and Seneca, 3.5 mol of review di Belle Arti NADPH per CO 2 ), than in feature articles its absence (3 mol of ATP and 2 mol of NADPH per for persuasive School, CO 2 ) . Thus it has been suggested that photorespiration is important for maintaining electron flow to prevent photoinhibition under stress . Photoinhibition occurs when the photochemical processes of photosynthesis do not have enough ADP or NADP to enable continued flow of energy through the electron transport system [71,72]. Electron flow becomes blocked as molecules within the electron transport chain become reduced. Then the electrons of the Photosystem II centre become highly excited as light energy continues to be absorbed; photoinhibition occurs as the excess light energy damages the D1 protein.
Photoinhibition is usually temporary because of the turnover rate of the repair mechanisms for the D1 protein. However, if other molecules within the photochemical system are permanently damaged by excess energy, chlorophyll becomes bleached, a process known as photo-oxidation. The risk of how to feature College photo-oxidation occurs under conditions such as salt and drought stresses, wherein internal leaf CO 2 concentration declines as stomata close to prevent water loss, electron consumption by photosynthesis is introduction for persuasive Warminster, reduced, and how to feature Seneca, over-reduction of the electron transport chain occurs . Photorespiration rises in plants under salt stress, as indicated by a higher CO 2 compensation point , higher light-to-dark ratio of CO 2 production , higher glycolate oxidase activity, and the formation of specific metabolites such as glycine, serine and custom, glycolate . CO 2 assimilation declines more quickly in write College heterozygous barley mutants with reduced levels of various photorespiratory enzymes than in the wild-type under moderate drought stress, demonstrating increasing control by photorespiratory enzymes on photosynthesis . Furthermore, Rubisco and photorespiratory enzyme activities are maintained in Casuarina equisetifolia L. Writing Scientific Articles School. under drought stress as the internal leaf CO 2 concentration decreases, suggesting that photorespiration is write Seneca College, likely to increase relative to photosynthesis . Photorespiratory glyoxylate exerts a feedback effect on photosynthesis in heterozygous barley mutated in International Eerde glutamine synthetase 2, ferrodoxin-glutamate synthase, GDC or serine:glyoxylate aminotransferase  (Figure 3). How To Write Feature College. Furthermore, glyoxylate can inhibit the activation of Rubisco in vitro , and writing scientific School, there is a negative relationship between glyoxylate content in the leaves of heterozygous glutamine synthetase 2 mutants and write feature articles Seneca, the activation state of Rubisco . Glyoxylate is also a reactive aldehyde that can modify lysine and arginine side chains, and form glyoxylated DNA adducts [80,81]. As indicated above, in vitro kinetic studies of recombinant proteins revealed that both cytosolic GLYR1 and plastidial GLYR2 catalyse the essentially irreversible, NADPH-based conversion of glyoxylate into glycolate, and can be regulated by custom essays King’s, the NADPH/NADP ratio [2–4].
Since the Michaelis constants for the two isoforms are apparently in the physiological range (4.5 and 34 ?M respectively), it is suggested that the enzymes function in the detoxification of glyoxylate in planta , and, in doing so, provide an alternative sink for excess electrons during the recycling of reducing equivalents. There is not yet any information on College expression of the GLYRs under photorespiratory conditions. HYPOTHETICAL ROLE FOR GLYRs DURING STRESS. Figure 3 summarizes the information discussed above with respect to custom written essays King’s Ely detoxification of write GABA-derived SSA and glycolate-derived glyoxylate in Arabidopsis by mechanisms involving the GLYR isoforms. In the absence of stress, flux through the GABA pathway should be relatively low, whereas photorespiratory flux would be considerable due to the favourable atmospheric CO 2 /O 2 ratio, and glyoxylate produced might contribute to coursework School Eerde GABA transamination to SSA. How To Feature Articles Seneca College. In the presence of stress, the cellular Ca 2+ or H + concentration could increase, thereby stimulating GAD activity and resulting in elevated GABA levels, and the oxygenation of ribulose bisphosphate would increase due to stomatal closure, probably resulting in glyoxylate accumulation. These responses would probably be associated with increases in NAD(P)H, which would limit the mitochondrial conversion of SSA into succinate via SSADH and glycine into serine via GDC. Based on the subcellular localization of the GABA shunt enzymes, it seems likely that GABA produced in the cytosol must be transported into the mitochondrion where it is custom, catabolized to how to write feature SSA by GABA-T activity. Writing. With the limitation of SSADH activity by elevated NADH concentration, SSA probably diffuses to the cytosol and perhaps the chloroplast, where it could be reduced to GHB via GLYR, a process facilitated by elevated NADPH levels. Similarly, some glyoxylate produced in how to the peroxisome probably diffuses to the cytosol and/or chloroplast to be detoxified by GLYR1 and/or GLYR2 respectively. Thus it is proposed that GLYR activity in plants serves to detoxify aldehydes that are typically generated in two distinct pathways, GABA metabolism and glycolate metabolism, and thereby contributes to redox balance.
The response of the pyridine nucleotides during stress could be attributed in large part to the biochemical control of NAD(H)K. In Arabidopsis , the three isoforms of NADK [cytosolic NADK1 and NAD(H)K3, and chloroplast NADK2] are essential for the phosphorylation of NAD(H) and have been linked to the plant stress response. Review Business Nuova Arti Milano. It is clear that NADK2 utilizes NAD only and that NADK3 prefers NADH, but the preference of NADK1 for NAD and NADH is less certain. How To Feature Articles Seneca College. Support, although preliminary, exists for activation by Ca 2+ /CaM of the NADK2 isoform only. Thus further research is required to provide a better picture of the relative importance and the biochemical control of the three NADK isoforms, particularly in view of the Ca 2+ /CaM activation of GAD, which is located in the cytosol. To test this scheme, various NAD(H)K expression mutants would be helpful in assessing the review business Nuova di Belle impact of altered redox balance on GABA metabolism and how to feature articles Seneca College, GLYR activity. Business Accademia. Additional experiments could be conducted with recombinant GLYR1 and GLYR2 to confirm whether the enzymes have the capacity to simultaneously reduce both glyoxylate and SSA, while recycling reducing equivalents. There is also a need to utilize/generate single and double knockouts of GLYR1 and GLYR2 in order to establish their physiological functions during stress. For example, the contribution of GLYR1 and feature articles Seneca, GLYR2 to the balance between energy input through photochemistry and energy utilization through metabolism (i.e. photostasis)  could be investigated by determining the levels of GABA/GHB, glyoxylate/glycolate and related amino and organic acids, as well as the redox (including the state of Photosystems) and energy balance and info International, photosynthetic efficiency. If the knockouts are more susceptible to stress than the wild-type, it could be concluded that the GLYR isoforms are involved in how to write articles College photostasis and prevent the accumulation of toxic aldehydes. At GLYR2 is localized to the chloroplast; however, its affinity for glyoxylate and SSA is 10-fold lower than that for GLYR1.
Typically, the over-reduction of the chloroplast electron transport chain by NADPH accumulation would probably be alleviated by GLYR2, but transgenic plants in which GLYR1 overexpression is coursework School, targeted to the cytosol or chloroplast could provide additional protection against photoinhibition and other stresses. If so, GLYR1 overexpression could become a strategy for engineering stress resistance in important crop plants. Work in the authors laboratory in this area is feature Seneca, supported by the Natural Sciences and Engineering Research Council of coursework Canada; and the Ontario Ministry of Agriculture Food and feature Seneca, Rural Affairs. We gratefully acknowledge the info International anonymous reviewers of the how to write articles College manuscript for many useful comments and scientific articles Pomfret School, suggestions. Abbreviations: AKR, aldo-keto reductase; CaM, calmodulin; GABA, ?-aminobutyrate; GABA-T, GABA transaminase; GAD, glutamate decarboxylase; GDC, glycine decarboxylase; GHB, ?-hydroxybutyrate; GLYR, glyoxylate reductase; HIBADH, hydroxyisobutyrate dehydrogenase; ICDH, isocitrate dehydrogenase; NADK, NAD kinase; ROS, reactive oxygen species; Rubisco, ribulose-1,5-bisphosphate carboxylase/oxygenase; SSA, succinic semialdehyde; SSADH, SSA dehydrogenase; TCA, tricarboxylic acid.
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How To Write A Feature Article | MJA
Worksheets in how to write Seneca VBA Coding and in Worksheet Formulas. Some miscellaneous SHEET coding. been transferred from my pathname, lastcell and buildtoc pages. Background information on Sheets (#intro) or if another workbook is involved, something like. Another way of creating your worksheet formula is to start by placing an research paper about equal sign on the formula bar to start a formula, then select a cell on the other sheet which could be in another open workbook. The formula is created for you.
Return to your original sheet and hit Enter. characters not allowed in write feature sheetnames : ? * [ ] Sheetnames may be up to Warminster 31 characters in length. Looping through a list of sheets Same idea but with array on write feature articles Seneca College, the For each statement: (Tom Ogilvy, 2006-02-24, programming) Create a list of Sheet Names from list in Column A, identified in Col B. Some Sheet related coding (#sheetcoding) Some CELL related coding and Row / Column related as well (#cellcoding) Number of pages to be printed (#pagenumbers) Thought I'd include this specifically from a posting by Eric Desart since there was some question as to coding. (all quotes in these formulas are double quotes) Note: Even though the syntax allows omitting a workbookname and sheetname, you should include it or you may be referring to the wrong sheet. (see Filename in a Cell, when the sheetname is omitted and the active sheet is used). Of possible interest: Display Page x of y in a cell, by Jan Karel Pieterse, in custom essays King’s his blog (Daily Dose of Excel), 2004-12-12, also see comments, and you can also get the row number of the page after a page break.
Don't copy a cell with ExceteExcel4Macro to another cell, copy from the formula bar instead (may crash otherwise, Jan Karel Pieterse) Build Table of Contents, similar listings, working with Hyperlinks Code to count all print pages from all worksheets from all open workbooks without activating that worksheetsworkbooks. Shailesh Shah, programming, 2002-10-30. For Excel 2007 users Shailesh has an add-on, “Exshail Classic Menu for Excel-2007 to make a usable menu from the less than useless Excel 2007 Ribbon. Check for Existence of a Sheet before creating (#existence) Check for write feature articles Seneca, Existence of introduction for persuasive Warminster a Sheet -- Worksheet Solution. Creating a series of sheets based on a master sheet (#master) Cycling through sheets looking for Seneca, specific value in cell A1 (#search) I think this is going to drive people crazy why not rename the sheets with the customer name or at least build a table of introduction essay Warminster contents with information.
See my buildtoc.htm page all you need to do is how to write articles Seneca College add content of cell A1 from each sheet. You will also find information there for sorting sheetnames. Scroll for position in window of active cell (#scroll) Display Cell A1 in scientific School upper left corner of write feature articles College sheet. Display Active cell in upper left corner of sheet. Navigations within a sheet based on another sheet (#navigation) Note both A23 and A24 refer back to essay writing Leysin American G23, so you cannot write a program directly reversible as where to feature go backwards in the macro from G23.
Navigation to a Sheetname (#moresheets) You can hold the custom essays shift key down click on the worksheet tab navigation arrows to shift over to the next set of worksheet tabs currently out of sight. Some Loops through Worksheets (#loopsheets) The LoopThru which resembles your original will reach a limit on characters in the msgbox -- limit varies by version. The LoopSheet will fail to how to write feature College show all sheets because due to limit or failure to be able to show another box after the first fills up the monitor window. More useful examples can be found in.
Build Table of Contents and optionally sort worksheets. Delete Active Sheet if name begins with sheet (#DeleteThisSheet) An exception for sheets with program code -- they will not be deleted nor will you be asked if you want to delete them. I have buttons to go through the sheet tabs, navigating to the Previous and Next described in BuildTOC, but you can use Ctrl+PageUP (to left) and custom written, Ctrl+PageDN (to right). The Toolbar icon for this macro is included on the toolbars page. Coding is with code/buildtoc.txt rather than code/sheets.txt might change the line above to the following to treat empty sheets the same as sheets beginning with Sheet:
Delete Empty Sheets (#delete_emptysheets) the second line is how you might type it in. the third line is how you might code the line for use in how to write articles Seneca INDIRECT. the fourth line invokes a user defined function (UDF) VBA goto subroutine/macro/function (#goto) ref. Application.Goto Reference:=Mid(ActiveCell.Formula, 2, i - 2) Coding a link address and friendly name using HYPERLINK Worksheet function (#hyperlink)
Until someone pointed out that you can use the writing Leysin School pound sign (#) By including the # the pathname can be left off, these work: in Excel 2000, and as I understand from Excel 97 and up, but these references are frozen within double quotes. See the next topic to take the references out from within the quotes. Hyperlink Worksheet References outside of quotes (#hyperlink_ref) To hyperlink and display the Cell in the B column same row of the Notes sheet. To hyperlink to a more descriptive longer note on the same row, but in the B column of the how to write College Notes worksheet. =HYPERLINK(#CELL(address,OFFSET( Notes !B1,ROW()-1,0)), Click for note) To hyperlink to paper about Warminster School the Note in Column B same row on Notes sheet, but display the short description in Column A of the Notes sheet. To refer to a cell in Column D (col 4) of the same sheet at a specific Row OFFSET to write articles the row. takes you down 41 rows to the first column, so the formula can be in any cell on for persuasive School, the row. The 4 indicates a relative address result, but it really doesn't matter if absolute or relative since it is how to articles Seneca College created during recalculation.
Will allow you to put the formula on research, any cell in the row , the link will take you 41 rows below and to the first column. Some alternatives to articles College Hyperlinks within a worksheet or within a workbook An Event macro such as double-click that asks for the first letter(s) of research paper about Warminster a lastname and then invokes the Match Worksheet Function to get you there. Feature Seneca College? The event macro example is essay American used with a name and address list sorted by lastname. An example of a Double-click (or right-click) event macro to goto another worksheet with the FollowHyperlink Method and specific cell using ActiveWorkbook.FollowHyperlink , there is how to write articles Seneca perhaps a better example on written, this page that goes to write feature articles another sheet with HELP information and provides the means of returning with the BACK button. CELL is a Volatile function, but worksheet volatile functions do not present the serious slowdowns that Volatile User Defined (VBA) Functions can have.
xlindex.htm is a sheet on my page, on my HD and on research Warminster, my website xlindex is also a sheet in my vlookup.xls file the how to write articles following examples show Column B what you see, Col C is the formula, and essay writing Leysin, C5-1 is the write feature articles College value in cell c5 on sheet xlindex. Values in RED are shown as constants if used in introduction for persuasive Warminster an actual formula referring to cells in the same workbook the formula is feature Seneca reduced to the formula shown immediately below -- no changes occur within double-quotes. Hyperlink to a MATCH (#hyperlinkmatch) You are looking for the value xyz (not case sensitive) in research about Warminster B1:B13. if you don't want to see #N/A then insert the failing part of the how to articles Seneca formula and the formula as follows: =IF(ISERROR(MATCH(xyz,B1:B13,0)), N/A, HYPERLINK(#ADDRESS(MATCH(xyz,B1:B13,0),2), xyz )) Don't ask me why the second formula hyperlink works but does not show up as a hyperlink, I don't know. You should look up ADDRESS Worksheet Function and the MATCH Worksheet Function in HELP for some details. Using HYPERLINK Worksheet Function to writing Pomfret School also pull in associated link (#vlookup)
On your sheet with VLOOKUP. Additional Examples with mailto. (#mailto) Using HYPERLINK Worksheet Function where workbook name can change (#internal) and use it as follows: A1: =HYPERLINK([hyperlink!$B$3 ]'Sheet One'!A1,Sheet One) But as updated in blue above, the following would be used within same sheet and would allow renaming/moving workbook without changing formulas. Feature Articles Seneca College? (works in XL97 and custom essays Ely, up) or within same workbook. Link Back to Sheet (#followhyperlink)
If you use a hyperlink or if in a macro you use FollowHyperlink Address you use hyperlinkaddress you can use the web BACK key (Alt+ArrowLeft) to return. Feature Seneca? You can install the buttons on your toolbar from tools, customize, commands, Web, and paper School, drag the two buttons BACK and FORWARD to how to articles Seneca College the toolbar. Invoke as a button on Worksheet. If your button is on the worksheet you can hyperlink to you help worksheet. I don't think you can do that you can assign a hyperlink directly to a toolbar button without having it invoke a macro instead. Invoke as a HYPERLINK Worksheet Function from the worksheet. =HYPERLINK(#helpme!E6,help) =HYPERLINK(#Help Me'!A1,help) Invoke as an object type hyperlink [Ctrl+K] Object hyperlinks (Ctrl+K) will =change automatically if you rename the coursework info worksheets, but can be burdomsome on older versions of Excel. (Pre 2000 on Win 98). Invoke link as a DoubleClick Event macro from the how to write worksheet.
Link to essay writing tricks Leysin American a Chart or Shape, faked by write feature Seneca College an Event Macro. Hyperlinks in Excel, Jon Peltier, some notes on hyperlinks including how to International fake a link to a chart (or other shapes). To Return to original worksheet: As mentioned at the beginning you can return with the Keyboard Shortcut ALT+ArrowLeft or with the Toolbar BACK button that can be installed on your toolbar as previously described. This topic was a little hard to decide whether to how to write College put on review Nuova Arti Milano, this webpage (sheets.htm), or on my Build Table of Contents webpage, or on Event Macros webpage as another example, or even perhaps the Shortcuts webpage because of the BACK button. Link Excel into HTML or MS Word (#linkexcel) for HTML as a href=c:tempmybook.xls#sheet1!c9worksheet 1/a Can't get the feature articles Seneca above HTML link to work in Firefox, let alone with a space in articles School the sheetname with Firefox. the following will SUM the feature Seneca College cells D3:G3 in each of the custom written essays King’s Ely sheets named in the cells L3:P3 (Dominic 2005-05-05, misc) -- You will get a #REF! error if any sheetname is missing or is invalid in the L3:P3 range. Some Worksheet Examples using INDIRECT (#indirect)
INDIRECT will not work for closed files. Extended one step further using the write articles College sheetname in cell A1: which now includes a single quote within double quotes around the sheetname specified in cell A1. You can simplify the coursework info School Eerde formulas by combining text together in all of the above (hope you can read it). Additional examples involving replication can be found on write, my Fill Handles page. INDIRECT to writing articles Pomfret refer to a cell in another workbook (actually used same workbook, can you tell?) Note cell A1 and E1 show a single quote but you must type in an additional single quote before the one you see.
Sheetnames are in Column C (cells C1:C50), The sheets that hae a model number of feature articles Seneca ModelA in their A3 cell are to review NABA, Nuova Accademia have the value of their H31 cells totalled in the formula. [Bob Phillips, .excel, 2006-05-31] Additional examples of INDIRECT can be found on my Build Table of Contents page. Referencing a Relative Worksheet Name in a Workbook (#relsheet) where MP1.xls is your other workbook. where 1 is the relative worksheet number.
where $A$2 is the cell in how to feature articles the other workbook's relative sheet 1. Open a workbook from writing Leysin School, named in write cells. List the names of the open workbooks. Generate Sheets with sheetnames named from current selection of cells. And if the macro is research School exited with any of how to write College these things in effect is there any way of telling what's wrong or just resetting it, I thought the following would put things back to normal, but after rereading the switches, I don't think it would change. Written Essays? I turned off macros and articles Seneca, fixed the interactive once, but more interested in written essays if you don't know what happened. C:Program FilesMicrosoft OfficeOfficeExcel.exe /regserver change path as needed or just use Excel.exe /regserver. 291288 - Startup Switches for Microsoft Excel (2002: 291288, 2000: 211481; 97: 159474) (SAFE Mode, Re-register/Reregister Excel extensions) Worksheet Tabs, and Linking (#hyp_ws_tabs)
Relative Hyperlinks and Hyperlink Base (#relative) But the use of the Hyperlink Base in write Seneca workbook properties can be used effectively, see thes postings by Harlan Grove. Scientific Articles? 2002-04-22 and this one to access the how to write feature Seneca College hyperlink property 2002-04-26. Missing Scrollbars or Worksheet Tabs (#missing) [x] Horizontal scroll bar. [x] Vertical scroll bar. [x] Sheet tabs [if still hidden then . Paper School? ] The horizontal scrollbar may have been dragged to far to the left, drag the Tab Split Box between the sheet navigation keys and horizontal scrollbar to the right to articles expose the Horizontal sheet tabs normally seen between the for persuasive essay navigation keys and the horizontal scrollbar. Things beginning with #-sign (#hash) Error messages: see help, index, also see Errors page. i.e. #N/A, #VALUE!, #REF!, #DIV/0!, #NUM!, #NAME?, or #NULL!, and for numbers or width ###### Cell Reference: To link to a cell or to a defined name, i.e. =HYPERLINK(#C5,C5) To link to a module: see Hyperlink to a code module, Andy Wiggins describes for each Excel version using the right click menu, equivalent to.
Master Sheet, creation (#mastersheet) Build Table of Contents, similar listings, working with Hyperlinks and less macro oriented variation Build TOC Another Approach. On this page see Looping through sheets Ron de Bruin has a web page specifically for a Master Sheet generation: Copy cells from each sheet in the workbook, topics include: Copy a range of write College each sheet, Copy the CurrentRegion of a cell of coursework International School each sheet, Copy the write feature Seneca College UsedRange of each sheet, Copy a row or rows of each sheet, Copy from row? till the last row with data, and Copy a column or columns from each sheet. Cells, Cells related coding. Build Table of Contents, listing of sheetnames and sheet codenames (see my site above). Written King’s? Check Existence of Sheet or sheet and WB and also, Chip Pearson, also see just for WS above (on this page). Create worksheets based on names in write feature College Column B, Robert Rosenberg, includes error recovery if sheet already exists. Collections , including Sheets collection, and references to info International hiding and unhiding worksheets (Hide and how to feature articles, Unhide sheets). Review NABA,? File Access with Visual Basic® for Applications, describes how to process files with the articles Seneca College file input/output (I/O) functions that Visual Basic for Applications provides -- information on using Sequentiai, Random access, and Binary. The examples show are for a .CSV type of file for Firstname, Lastname, etc. Warminster School? Grouping worksheets, to make manual changes to how to write feature articles several sheets at one time.
Grouping worksheets via an Array, which could be doing more refined selections, Jim Rech, 2003-08-15, programming Importing Data into Excel (.CSV, .html) import.htm -- rough sketch. Info? Menu, Worksheets into a, Chip Pearson, see entire thread. How To Write Articles College? Move, Copy, Replace, Populate Optimizing VBA, Chip Pearson Pathname in headings, footers, and cells Print tips for Excel, Ron de Bruin Referencing Worksheets From Formulas (C. Pearson), VBA procedure that you can use to refer to the first, next, previous, or last worksheet in written Ely a workbook, called directly from worksheet cells. Summary of Methods for Navigating between worksheets that you will find here in My Excel Pages.
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3 Tragic Misconceptions About Proposal Cover Letters. Cover letters are one of the most important pieces of a proposal. How To Write Feature Articles Seneca. Unfortunately, 99% of the coursework Eerde cover letters I#8217;ve seen during my Brutally Honest Proposal Critiques are cr*p. Articles Seneca. The problem stems from common misconceptions about the writing scientific School cover letter#8217;s place in the current proposal environment. Lets look at three misconceptions that can lead to epic cover letter failure. How To. Misconception #1: People Want To Read Your Cover Letter. Recently, I was at a prebid meeting at PASSHE, which is responsible for dorm construction for Pennsylvania#8217;s state university system. The Director of Construction made it plain. #8220;Don#8217;t include a cover letter in your proposal.
It#8217;s not going to convince us. So, don#8217;t waste your time.#8221; Why would he say that? He said that because he#8217;s seen hundreds, if not thousands, of cover letters and they all said nothing. There was nothing of value in those cover letters. Naturally, when he sees a cover letter, we have trained his brain to skip over it. Custom Essays King’s Ely. If you think someone is likely to read your cover letter, you are insane. Instead, you have to give people a reason to how to write feature articles, read your cover letter.
You have to get their attention and then create a #8220;slippery slope#8221; that will keep them reading. I give an in-depth tutorial on creating a slippery slope in Proposal Development Secrets. Many of King’s Ely, you have already read the book. How To Write Feature Articles College. I won#8217;t rehash all of that. Introduction For Persuasive Essay Warminster School. But remember, you have to start off by creating a #8220;knowledge gap.#8221; You create a knowledge gap by highlighting a gap between what the reader knows and what the articles Seneca reader wants to know. Let#8217;s say you are proposing to a client that knows your firm pretty well.
You might use, #8220;3 Things You Might Not Know About ABC Architects.#8221; Those three things better be something they don#8217;t know and also highlight what they would be missing out on if they hire someone else for writing scientific, this project. You#8217;ll notice I use this knowledge gap technique with a lot of my titles. That#8217;s why my open rate is so much higher than the industry average. I give people a reason to read, then deliver. Obviously, it#8217;s a technique I#8217;ve also used quite successfully in proposals to clients all over the country. Feature College. Misconception #2: You Can Use A Template. People always get angry at me when they ask for an example of writing Pomfret School, a great cover letter for them to articles College, use. That#8217;s because I tell them such a thing does not exist.
A great cover letter to the Department of review Accademia Arti, Veterans Affairs is not going to be a great cover letter to the University of Pennsylvania Health System. Each cover letter has to speak to the specific client and how to write feature articles Seneca College, project. Unfortunately, that means, gasp, each cover letter will have to be written from scratch. There is no way around it. Well, actually there is. Write a terrible cover letter they will simply skip over. Research Paper About Warminster School. But that won#8217;t help you win. Misconception #3: Throw Everything At Them and how to feature Seneca College, See What Sticks.
This misconception leads to written King’s Ely, long cover letters. Here#8217;s the problem. If you had received a three-page letter in 1973, the likelihood that you would have read all three pages would have been very high. But we live in a different world. When was the last time you read, word for word, a three-page letter? Even better yet, when was the last time you even received a three-page letter? I#8217;m sorry to say that if my own grandmother sent me a three-page letter, she#8217;d be pushing her luck.
I couldn#8217;t promise I#8217;d read ever word. Busy people just don#8217;t read long cover letters. How To Seneca. Anything longer than a page is pushing it. When my boss sees a two-page cover letter from me, his initial reaction is, #8220;There is something up. Why is this so long?#8221; Two-page cover letters are sometimes unavoidable. But three-page cover letters are always avoidable. Every word needs to earn its place on the cover letter. If it doesn#8217;t speak specifically to this client, to this project#8230;it hasn#8217;t earned its place. As much as it pains you, leave it out. Don#8217;t let these misconceptions plague your cover letters. Follow my advice and you#8217;ll get an opportunity to make a powerful pitch to your clients.
If you want to learn more about the slippery slope, I just released an unrestricted PDF of Proposal Development Secrets. Writing Scientific School. If you liked this article, please subscribe below or on the right side of the homepage. If you want to give us your thoughts on this issue, please leave a comment below. Great post! Misconception #2 is spot on! It drives me nuts how many times I see the same generic letter over how to articles College, and over#8230; Also, this template letter usually has everything in it (Misconception #3). Thank you for writing tricks School, the comment. How To Write Seneca. I love reading cover letters that say #8220;We#8217;re excited to be submitting on project XYZ#8230;#8221; Yes I can tell you are excited to Ely, be submitting#8230; (in my best Ben Stein voice) That#8217;s why your cover letter states it. Our it states you are passionate about the project. People who are passionate don#8217;t have to write they are passionate it shows in how to write feature articles Seneca College everything they do or write. Research Paper Warminster School. Another great post worth passing along!
Good reminders. This is similar advice to an SMPS speaker I saw at a regional conference. How To Write Articles College. Here is where I struggle though. Any advice on tricks Leysin School helping BD staff get good project-specific, client-specific info to write articles College, pack into proposals? Or to help find out what we offer that#8217;s actually different? I#8217;m the marketer who doesn#8217;t interact with clients. All my info comes from BD and if I don#8217;t get good info from them the entire proposal, not just the cover letter, reflects that. I feel there has to custom written King’s Ely, be a better way because otherwise it seems impossible to do what you say. Thank you for the comment.
I#8217;ll address this problem in a future post. Keep reading. How To Feature College. I do talk a little bit about this in my book. Pick that up if you haven#8217;t yet. Here#8217;s the link. Cover letters are a waste of everyone#8217;s time. I wouldn#8217;t go that far. I certainly don#8217;t think they are a waste of time. Custom Essays Ely. Not if you do it right. I was once told by a US Army Corps of Engineers that when a proposal has a bound-in cover letter, the write feature Seneca College secretary rips the letter out of each copy before distributing them to the screening committee. That#8217;s a good story.
I believe it. Years ago, I was on a debrief call where the Army Corps admitted to custom written, not reading most of our proposal. How To Write Feature. I#8217;m the only proposal writer at coursework info School, an engineering firm with nearly 20 project managers. All but a very small minority consistently rely on how to write articles at least one of these three misconceptions for coursework School Eerde, every single cover letter. thanks Matt. Write Articles. You might start by forwarding them the link to this article. Tricks American. Sometimes it helps to write, hear it from an tricks Leysin outside source.
I#8217;ve made everyone of those mistakes in cover letters, multiple times. Feature Articles Seneca. As usual, you are spot on. Thanks Matt. Essay Warminster. Thank you for commenting. I didn#8217;t even know you read the site. #128578; I#8217;ll shoot you an email. Everything you are saying about the how to write articles College old way about essays Ely writing cover letters is true.
I hate having to follow what the uncreative people wrote before. Even though we win most of our bids, I would still like to write fresh and creative cover letters and proposals. Thank you for write feature College, commenting. Cover letters are just a piece of the puzzle. I#8217;m sure many successful firms have terrible cover letters. Yet, they still win lots of contracts. Research Warminster. But even if you have a winning record, getting even better will help protect your position. It sounds like you agree. Great post, Matt. I have been pushing people toward more concise and how to College, engaging letters for years and will keep fighting the custom King’s Ely good fight.
When I have just a few minutes to write feature articles Seneca College, #8216;peer review#8217; a colleague#8217;s proposal, I always go to the letter first. Introduction Essay. Whether read, skimmed, or glanced at, the letter remains part of the feature articles client#8217;s first impression. Thank you for research paper about Warminster, your comment Jeff. Keep fighting the how to articles College good fight. This is a great piece, Matt. To go along with it, I remember seeing (long ago) a list of the phrases we all used in writing scientific articles Pomfret School cover letters which one should never use. For example, the write feature articles Seneca College first item on the list was the traditional opening, #8220;We are pleased to present.#8221; The explanation was that if you weren#8217;t pleased to present, you wouldn#8217;t have presented at all, so these words were a waste of introduction for persuasive School, space. If someone could find that list again and feature, share it, maybe we#8217;d all learn even more about cover letters than you taught above. Was it words not to scientific Pomfret, use in your proposal?
If so, I have that document. When will we be able to turn off the broken record and stop having to repeat ourselves when we tell our folks in the field #8220;quality, not quantity. #8221; the Proposal Writer lamented, tapping dejectedly at write feature Seneca College, her keyboard. Cover letters can be quite valuable if the writer uses them appropriately, e.g., as a way to connect personally/remind the prospect of a recent meeting and to segue into custom Ely, the proposal with a friendly but professional recap of the reader#8217;s needs and objectives (if the salesperson was lucky enough to have had an actual meeting or even phone call with the how to feature prospect ahead of time). Nice choice of graphic. I like the oh-so-subtle hidden #8220;hurdle#8221; concept as a backdrop for the massive wipeout. Thank you for the comment. Essays Ely. Interesting reading I was in a proposal training where it was stated in how to write Seneca today#8217;s world it is best practice to not include a cover letter as it has no value add. I#8217;m old school and agree it is just good business practice and one should take the opportunity to make it of value. Thank you for the comment. Matt this is great. Thanks for essay writing Leysin School, giving voice to my frustration with template letters.
Thank you for commenting. Great post! I like your idea for #8220;3 things you may not know about our company#8221;. [#8230;] Here#8217;s a great piece by Matt Handal about how to feature articles College proposal cover letters. [#8230;] Get The FREE Proposal Writing Crash Course! Proposal Workshop (Austin, TX) How to Get Meetings with the Most Unreachable Potential Clients (Free Webinar For SDA Members) Proposal Development Secrets (Free Webinar) How to writing articles Pomfret, Get Meetings with the Most Unreachable Potential Clients (Free Webinar) How Much Traffic Should Your Website Really Get? Why Mistakes In Proposals Don#8217;t Matter…and What Does. The Ultimate SF330 Guide (with Template) A/E Marketers Are Not Your Gal Friday. How To Feature Seneca. What I#8217;ve Learned From Using Zurmo As My CRM.
The Purpose of Graphics and Images in writing scientific School a Proposal…It#8217;s Not What You Think. AEC Marketing Jobs Marketing and Business Development Jobs With Architecture, Engineering, and Construction Firms Business Development Manager Job Description A new site about business development CofeBuz Tim Klabunde#8217;s tips on becoming a better networker. Construction Marketing Ideas Mark Buchshon#8217;s daily blog for the construction marketer. Delay Damages A website all about how to feature College construction delay damages. Ediscovery An entire site about ediscovery How To Write A Proposal A website dedicated to explaining proposal writing to those who have little to no experience doing so. Marketers Take Flight Career advice for A/E/C marketers. My Business Letter The internet#8217;s more comprehensive website on essay tricks Leysin School the business letter. Hi. I'm Matt Handal. College. And this is my site Help Everybody Everyday.
I provide helpful advice to marketers in the architecture, engineering, and NABA, di Belle Arti, construction world.
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